Perplexing article.
I visited the WAP (Niger+Benin+Burkina) and I reserve my opinion.
I have to check the credencials of the authors. So far the best specialist for lion is Philippe Chardonnet, a veterinarian and hunter, who is an undiscuted expert in lion and our best defender against the banning of lion hunting. His opinion and research projects are yet accepted by the CITES.